How are the PCIe lanes of the JUNO system connected

Hi,

the documentation states:

• 4 lane Gen 2.0 PCI-Express (Juno r1 and r2 only)

The JUNO system has 4 PCIe connectors:

     x16, x8, x4, x4

Does every connector have 4 independent lanes (so all together we could use 4 cards with 4 lanes each at the same time)

or are the 4 lanes shared so when we are using 4 cards only one lane is available per card?

Thanks,

           Martin